o:22023
Ontogenetska, histološka, semiohemijska i etološka istraživanja vrste Pachyiulus hungaricus (Karsch, 1881) (Diplopoda, Julida, Julidae)
doktorska disertacija
Ontogenetic, histological, semiochemical and ethological research of Pachyiulus hungaricus (Karsch, 1881) (Diplopoda, Julida, Julidae) : doctoral dissertation
sr
U ovoj tezi predstavljeni su rezultati istraţivanja razliĉitih aspekata biologije
diplopode Pachyiulus hungaricus (Karsch, 1881): kasnog perioda postembrionalnog
razvića, histološke graĊe odbrambenih ţlezda, sastava i antimikrobnog dejstva
odbrambenog sekreta i reproduktivnog ponašanja.
Euanamorfno razviće je prouĉavano kroz stadijume VII-XII, odnosno, kroz period
kasne postembriogeneze. Definisanje stadijuma izvedeno je na osnovu broja nizova ocela,
dok vrednosti analiziranih morfoloških osobina imaju dopunski informacioni znaĉaj.
Adultne jedinke sa funkcionalnim gonopodama i funkcionalnim vulvama uoĉene su na
stadijumu XI.
Histološka graĊa odbrambenih ţlezda je analizirana kod adultnih jedinki.
Komponente odbrambenog sekreta su lokalizovane u sekretornim granulama, koje se, na
osnovu odreĊenih morfoloških osobina mogu svrstati u nekoliko tipova. Rezultati
ultrastrukturne analize ukazuju da se sekret kanalima doprema u lumen rezervoara, što
odgovara merokrinom tipu sekrecije.
Dominantna jedinjenja u sekretu P. hungaricus su hinoni, dok su nehinonska
jedinjenja u vidu pentil i heksil estara zasićenih i nezasićenih masnih kiselina dugog niza.
Sekret je ispoljio inhibitorno dejstvo na rast gljiva i bakterija, kao i baktericidno i
fungicidno dejstvo.
S obzirom da u privlaĉenju partnera mogu da deluju i hemijski stimulusi, prouĉavan
je aspekat reproduktivnog ponašanja, prekopulatorno ponašanje. Opisano je nekoliko
sekvenci ponašanja u testu nazvanom „arena za parenje“, a zatim u testovima izbora
muţjaka i izbora ţenki. Nekoliko morfoloških osobina jedinki koje su se parile i onih koje
nisu imale uspeha u parenju, analizirano je metodama tradicionalne i geometrijske
morfometrije. Rezultati su pokazali da je osobina oblika nogu podloţna prekopulatornoj
seksualnoj selekciji.
In this thesis, the results of research on various aspects of biology of diplopod
Pachyiulus hungaricus (Karsch, 1881) are presented: late period of postembryonic
development, histological structure of defensive glands, composition and antimicrobial
effect of defensive secretion and reproductive behaviour.
Stadia VII-XII, which belong to the late period of euanamorphic development, were
included in the analysis. Counting rows of ocelli was the primary method for differentiating
between stadia, while the other analysed morphological traits are of supplementary
informational significance. Adults possess functional gonopods and functional vulvae at the
stadium XI.
Histological structure of defensive glands was investigated in adults. Components of
defensive secretion are localized in secretory granules, which could be classified into
several types, based on their morphology. Results of ultrastructural analysis indicate that
ducts transport the secretion into reservoir lumen, which corresponds to the merocrine
secretion mode.
Dominant compounds in the secretion of P. hungaricus are quinones, and nonquinonic
compounds are pentyl and hexyl esters of long-chain fatty acids. The secretion
had inhibitory effect on the growth of tested bacteria and fungi, as well as bactericidal and
fungicidal effect.
Considering that chemical stimuli could be important in the mate attraction, we
studied the aspect of reproductive behaviour, precopulatory behaviour. Several sequences
of the behaviour were described during mating arena test and then in male and female
choice tests. Also, several morphological traits of mated and nonmated individuals were
analysed using traditional and geometric morphometrics. The results indicate that the leg
shape is subjected to precopulatory sexual selection.
Biologija - Biologija razvića životinja / Biology - Biology of Animal Development Datum odbrane: 27.12.2019.
Pachyiulus hungaricus, Diplopoda, euanamorfoza, morfološka varijabilnost,odbrambene ţlezde, sekretorne ćelije, odbrambeni sekret, antimikrobno dejstvo, uspeh u parenju
Pachyiulus hungaricus, Diplopoda, euanamorphosis, morphological variabilitydefensive glands, secretory cells, defensive secretion, antimicrobial effect, mating success
591.5:595.61(043.3)
yes
1025241010
91552101
7446
91552100
1025241010
2020-04-09T14:50:18.053Z
45
no
46
mentor
Zvezdana, 1989-, 32836199
Jovanović
2019
63
mentor
Slobodan, 1963-, 12628071
Makarov
2019
63
mentor
Luka, 1965-, 12692071
Lučić
2019
63
član komisije
Aleksandra, 1964-, 12662631
Korać
2019
63
član komisije
Sofija, 1967-, 16587111
Pavković-Lučić
2019
63
član komisije
Ljubodrag, 1978-, 13872487
Vujisić
2019
141 list
8842917
http://phaidrabg.bg.ac.rs/o:22023
no
yes
4
2020-04-09T14:50:18.320Z
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Pachyiulus hungaricus, Diplopoda, euanamorfoza, morfološka varijabilnost,odbrambene ţlezde, sekretorne ćelije, odbrambeni sekret, antimikrobno dejstvo, uspeh u parenju
Pachyiulus hungaricus, Diplopoda, euanamorphosis, morphological variabilitydefensive glands, secretory cells, defensive secretion, antimicrobial effect, mating success
591.5:595.61(043.3)
1738
11A02
2019